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Approximate age:3,600,000 years old=average of minimum (3.6 Ma) and maximum (3.6 Ma)
Date based on: K-Ar See map of sites with similar ages
Dating reference: Deino, A. L., Scott, G. R., Saylor, B., Alene, M., Angelini, J. D., & Haile-Selassie, Y. (2009). 40Ar/39Ar dating, paleomagnetism, and tephrochemistry of Pliocene strata of the hominid-bearing Woranso-Mille area, west-central Afar Rift, Ethiopia. Journal of human evolution.
Hominids: Australopithecus afarensis
Archaeology:
Important specimens: KSD-VP-1/1 partial skeleton, MSD-VP-5/16 mandible
Notes:
Location: Afar Rift, Ethiopia
Map location/coordinates reference: Haile-Selassie, Y. (2010). Phylogeny of early Australopithecus: new fossil evidence from the Woranso-Mille (central Afar, Ethiopia). Phil. Trans. R. Soc. B 365(1556), 3323-3331
Haile-Selassie, Y., Latimer, B. M., Alene, M., Deino, A. L., Gibert, L., Melillo, S. M., et al. An early Australopithecus afarensis postcranium from Woranso-Mille, Ethiopia. Proceedings of the National Academy of Sciences, 107(27), 12121.
Key references: Haile-Selassie, Y. (2010). Phylogeny of early Australopithecus: new fossil evidence from the Woranso-Mille (central Afar, Ethiopia). Phil. Trans. R. Soc. B 365(1556), 3323-3331
Abstract
The earliest evidence of Australopithecus goes back to ca 4.2 Ma with the first recorded appearance of Australopithecus anamensis at Kanapoi, Kenya. Australopithecus afarensis is well documented between 3.6 and 3.0 Ma mainly from deposits at Laetoli (Tanzania) and Hadar (Ethiopia). The phylogenetic relationship of these two species is hypothesized as ancestordescendant. However, the lack of fossil evidence from the time between 3.6 and 3.9 Ma has been one of its weakest points. Recent fieldwork in the Woranso-Mille study area in the Afar region of Ethiopia has yielded fossil hominids dated between 3.6 and 3.8 Ma. These new fossils play a significant role in testing the proposed relationship between Au. anamensis and Au. afarensis. The Woranso-Mille hominids (3.63.8 Ma) show a mosaic of primitive, predominantly Au. anamensis-like, and some derived (Au. afarensis-like) dentognathic features. Furthermore, they show that, as currently known, there are no discrete and functionally significant anatomical differences between Au. anamensis and Au. afarensis. Based on the currently available evidence, it appears that there is no compelling evidence to falsify the hypothesis of chronospecies pair or ancestordescendant relationship between Au. anamensis and Au. afarensis. Most importantly, however, the temporally and morphologically intermediate Woranso-Mille hominids indicate that the species names Au. afarensis and Au. anamensis do not refer to two real species, but rather to earlier and later representatives of a single phyletically evolving lineage. However, if retaining these two names is necessary for communication purposes, the Woranso-Mille hominids are best referred to as Au. anamensis based on new dentognathic evidence.
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